Columns

These settings apply to Folder Columns.

Create new columns without animating: When it's off and you click on a folder in the Current Folder Column, the new column slides in from the right instead of appearing instantaneously. The sliding speed is determined by the global animation duration.
Column Pane alignment: Sets the alignment of folders in the Folder Panes individually for Standard and Columns layouts. You may not notice the effect of this setting unless the columns are reasonably wide.

Interaction

This refers to the interaction between cursor and columns. One Commander's default is to make Folder Columns expand when the cursor enters them and contract when it leaves. Some users like this because it makes it easier to view columns when a great many are displayed (i.e. when the Current Folder is a long way from the drive root); others don't.
Expand Column when cursor enters it: For each Layout, turns on this behavior.

Advanced Settings

Don't be put off by the "Advanced" label; it simply indicates that these are settings that many users won't be interested in using. They adjust things like how quickly columns expand when the cursor enters them (both in milliseconds), how long the expansion takes, what width of column One Commander should apply the expansion to, and a number of settings for column widths, and they're self-explanatory. They're worth experimenting with: you may find that they improve One Commander's usability considerably. For example, shortening Expand delay and Expand duration (say, to 400 ms each) makes the column expansion and contraction much more responsive. If the delays are too short, the columns will expand when the cursor is moving quickly over them, which could get annoying.
If you come up with a mixture of settings that you really like, please, as the note says at the bottom of the tab, share your findings with [email protected]

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Backlinks:
2. Quick introductory guide:4. Folder columns
3. Full reference guide:Folder Columns and Navigation Pane
3. Full reference guide:Settings:General